physical, chemical, or biological perturbations to a system that are either (a) foreign to that system or (b) natural to the system but applied at an excessive [or deficient] level (Barrett et al. 1976:192). Stressors cause significant changes in the ecological components, patterns and processes in natural systems. Examples include water withdrawal, pesticide use, timber harvesting, traffic emissions, stream acidification, trampling, poaching, land-use change, and air pollution.